Ross Brawn is expected to leave his position as team principal of Mercedes following their recruitment of Paddy Lowe from McLaren.
Lowe is to run the team along with new Mercedes sports boss Toto Wolff from 2014 but he will be employed by McLaren until then on 'gardening leave'.
Brawn will stay on for now but sources close to Mercedes say he will not continue in the long term.
McLaren have promoted engineering director Tim Goss.
Mercedes are refusing to comment.
He will fulfil a role senior to technical director, running the sporting and technical aspects of the team, while Wolff fulfils other senior responsibilities, such as political and commercial.
That set-up would appear to leave no room for Brawn in his current role.
Brawn said last month in the wake of BBC Sport's revelation: "Paddy is not coming. I want to see how things go before I make a final long-term commitment.
"If I choose to leave the team then Paddy will come. We've got a plan. We have a fall-back plan. It's as simple as that."
In an interview with the official Formula 1 website on Monday, Brawn said: "I am responsible for the sporting side; running the team on an operational level. If we can maintain that then there will not be a problem."
Lowe's arrival at Mercedes, which has been put in place by Wolff and the team's non-executive director Niki Lauda, also raises questions about the senior engineering team at Mercedes.
Including Brawn, they have four men who have been previously technical directors at other teams - technical director Bob Bell, technology director Geoff Willis and engineering director Aldo Costa.
BBC Sport understands that Wolff and Lowe are analysing the current structure and will make changes if they perceive there to be any weak links.
Lowe's move means he will join forces again with Lewis Hamilton, who left McLaren at the end of last season to join Mercedes.
It is the first sign of the ruthlessness with which Mercedes are pursuing success in F1. Earlier this month, Wolff made it clear he would do whatever it took to win the championship and emphasised that failure was not an option.
Brawn is the man behind all seven of Michael Schumacher's world titles with Benetton and Ferrari but he has been less successful in his current position.
The team won the drivers' and constructors' championships with Jenson Button in 2009 under Brawn's name, but that was partly because of the unique circumstances of the controversial "double diffuser" technical advantage and the huge financial input of former owners Honda before they pulled out at the end of 2008.
They have won only one race since Mercedes took them over for the 2010 season.
In a statement released on Monday McLaren's team principal Martin Whitmarsh said of Goss's appointment: "I'd like to offer my congratulations to Tim on his new position, which, after more than 20 years of exceptional service for McLaren, he richly deserves."
On Lowe's departure, Whitmarsh added: "Paddy [Lowe] will be performing a different role within McLaren until the end of the year. He's been a good and successful F1 technical director, and we wish him well when he embarks on a fresh challenge in 2014."